Dan graduated from the University of Bristol in 1997. He moved to the School of Veterinary Science, University of Liverpool in 2002. He became a Diplomate of the European College of Veterinary Internal Medicine in 2007. In 2010 he completed PhD studies in the field of molecular physiology, researching the regulation of intestinal glucose transporters in dogs and cats, and their relevance to diabetes mellitus and obesity. He rejoined the Medicine team at the Small Animal Teaching Hospital, University of Liverpool in 2011 and is currently Senior Lecturer in Internal Medicine and Head of Small Animal Medicine there.